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In practice, large empirical datasets sx natural populations incorporating stage- and sex-specific vital rates are uncommon 25 — Furthermore, males and females often have different behaviors or ecological niches 28which can make one or the other easier to detect 9 Fortunately, these sources of sampling bias can be accounted for using mark—recapture methods Here, we investigate the demographic origins of ASR bias in a polygamous bird using 7 years of individual-based sex- and stage-specific life history data. Polygamous species have a special significance in sex ratio studies because they are predicted to be at higher risk of extinction 1 We studied a small ground-nesting shorebird, the snowy plover [Charadrius nivosus 31 ], which is endangered in parts of its Nearctic range and has a sequentially polygamous mating system 32 Using a two-sex matrix model, we show that the ASR of this species is substantially more male-biased than previously reported Sex differences in chick and juvenile survival contribute most to ASR bias, suggesting that ASR variation is particularly susceptible to factors that influence early life history stages.

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